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pcie gen 4 in gen 3 slot
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4-Pin Power Connector up to 5 Gbps Speed PCIe Card · 6 out of 5 Tp-Link Tx401 10 Gbps Wireless Gigabit Pci Express Network AdapterPcie Slot

The following image shows the PCI connections of the motherboard used in DuroPC's IMB540-4 Notice 'SLOT 13' are PCIe x16 slots but only Slot 3 has an x8

pcie 3 slot PCI Express® also known as PCIe or PCIe Gen 4 is the fourth generation of Peripheral Component Interconnect Express

pcie slot pinout 2 into a normal PCIe x1 slot, and leaves the remaining 3 lanes unconnected Gen 4 drive at full speed in a PCIe slot: WD SN850 1TB in x4